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The DN entries you specify in the User DN for Query Authentication and the Base Search DN field are, in effect, search paths in an LDAP structure.
Consider the DN examples used in Administering AE Services access to Active Directory:
User DN for Query Authentication
cn=Pat Brown,cn=sertech,cn=services,cn=users,dc=example,dc=com
Base Search DN
cn=users,dc=example,dc=com
Both DNs are unique, but the User DN for Query Authentication is more specific than the Base Search DN.